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clock-iconPUBLISHEDFebruary 12, 2025
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New Fish Species With Stripey “Face Paint” Named After Iconic Studio Ghibli Character

Its scientific name is a nod to Princess Mononoke’s San.

Holly Large headshot

Holly Large

Holly Large headshot

Holly Large

Copy Editor & Staff Writer

Holly has a degree in Medical Biochemistry from the University of Leicester. Her scientific interests include genomics, personalized medicine, and bioethics.

Copy Editor & Staff Writer

Holly has a degree in Medical Biochemistry from the University of Leicester. Her scientific interests include genomics, personalized medicine, and bioethics.View full profile

Holly has a degree in Medical Biochemistry from the University of Leicester. Her scientific interests include genomics, personalized medicine, and bioethics.

View full profile
EditedbyFrancesca Benson
Francesca Benson headshot

Francesca Benson

Copy Editor and Staff Writer

Francesca has an MSci in Biochemistry from the University of Birmingham.

photograph of Branchiostegus sanae, a fish with red and white vertical stripes under its eye, overlayed on an illustration of the sky done in a Studio Ghibli-esque style

Just to clarify, Branchiostegus sanae definitely doesn't fly.

Image credit: Huang et al., ZooKeys 2025 (CC BY 4.0)/Creative Kiosk/Shutterstock.com; modified by IFLScience


Scientists have discovered a brand-new species of deep sea-dwelling fish, sporting a unique facial feature that led the team to name it after a character from the animated film Princess Mononoke, a cult classic produced by Studio Ghibli.

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The fish – which belongs to the family Branchiostegidae, commonly known as deepwater tilefish – has two vertical stripes under its eyes, one white and one red. These stripes reminded the researchers of San, a central character in Princess Mononoke, who also has red striped facial markings.

But it’s not just facial features that led the team to name the new species Branchiostegus sanae – the connection goes deeper.

“In Princess Mononoke, San is a young woman raised by wolves after being abandoned by her human parents. She sees herself as a part of the forest and fights to protect it,” explained lead author Haochen Huang in a statement. “The film delves into the complex relationship between humans and nature, promoting a message of harmonious coexistence between the two: something we hope to echo through this naming.”

The new species also wasn’t discovered in what could be assumed as the “typical” way. Rather than a team of researchers scouring the depths of the ocean, B. sanae was instead spotted when Huang and colleagues were having a scroll through online seafood markets back in 2021. And just like that, there’s a new excuse for the next time your boss catches you on your phone during work hours – you could be about to discover a new species!

Branchiostegus sanae at a seafood market.
Branchiostegus sanae at a seafood market.
Image credit: Jiangyuan Chen

It’s not unusual to find a deepwater tilefish in such a market – they’re a popular type of seafood in East and Southeast Asia – but it was B. sanae’s facial markings that caused it to stand out among the rest, and it turned out to be a particularly unique find.

“Finding a new species in this group is a rare and fortunate event, especially one as distinctive as Branchiostegus sanae,” said Huang. Indeed, there are just 30 species in the deepwater tilefish family, with B. sanae only the second to be described in the last decade. It’s also believed to be the only tilefish species known to live in the South China Sea that has vertical stripes on its body. 

By working with Chinese fishermen – who refer to B. sanae as “ghost horsehead fish” – the researchers were able to determine the specimens were collected from an area between the Xisha Islands and Hainan Island. 

Deepwater tilefish are typically found in regions on the edges of oceanic and continental plates, where they hang out on the ocean floor at depths between 20 to 200 meters (66 to 656 feet).

The study is published in the journal ZooKeys.


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